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On the top, protein expression in current human tissue, based on all annotated cell types, is reported with the units not detected (n), low (l), medium (m) and high (h). Underneath, protein expression in each annotated cell type are reported using the same units.
Protein expression data is based on knowledge-based annotation. For genes where more than one antibody has been used, a collective score is set.
If knowledge-based annotation could not be performed for a gene, no data is displayed here. View antibody staining data further down this page.
A summary of mRNA expression data available for current tissue based on several datasets. The mRNA expression levels in human tissues are based on RNA-seq data generated by the Human Protein Atlas (HPA), Genotype-Tissue Expression (GTEx) portal and CAGE data generated by the FANTOM5 consortium. Consensus normalized expression levels for human tissues was created by combining the data from HPA and GTEx datasets.
The mRNA expression levels in pig are based on RNA-seq data generated by the Human Protein Atlas (HPA), and for mouse, HPA data and in situ hybridization generated by the Allen brain atlas are reported.
Scroll down to view mRNA expression data in more detail.
RNA-Seq data generated by the Genotype-Tissue Expression (GTEx) project from human tissues is reported as mean nTPM. More information can be found on the GTEx portal.
Distribution across the dataset is visualized with box plots, shown as median and 25th and 75th percentiles. Points are displayed as outliers if they are above or below 1.5 times the interquartile range. nTPM values of the individual samples are presented next to the box plot.
Brain - Nucleus accumbens (basal ganglia) Average nTPM: 1152.8
Samples: 246
Max nTPM: 4348.9
Min nTPM: 413.0
Std nTPM: 405.1
Median nTPM: 1079.3
GTEX-147F4-0011-R6b-SM-5S2UP
50-59 years, male
4348.9
GTEX-14753-0011-R6b-SM-5S2UD
60-69 years, male
3026.4
GTEX-1EKGG-0011-R6a-SM-CL538
60-69 years, male
2890.2
GTEX-1LC47-0011-R6b-SM-EZ6MI
60-69 years, male
2733.3
GTEX-11DYG-0011-R6a-SM-DO936
60-69 years, male
2453.4
GTEX-11DZ1-0011-R6b-SM-DNZZG
50-59 years, male
2171.9
GTEX-1LVAO-0011-R6b-SM-EZ6MX
60-69 years, male
2122.1
GTEX-1H3O1-0011-R6b-SM-CL54U
60-69 years, male
2042.8
GTEX-X4XY-0011-R6B-SM-46MVB
60-69 years, male
2035.0
GTEX-13FHO-0011-R6b-SM-5LZX1
60-69 years, male
2032.7
GTEX-ZV68-0011-R6a-SM-51MSR
50-59 years, female
1796.7
GTEX-11PRG-0011-R6b-SM-DO923
50-59 years, male
1791.9
GTEX-1RLM8-0011-R6a-SM-EYYXS
50-59 years, male
1724.7
GTEX-1HBPH-0011-R6a-SM-CKZOV
40-49 years, female
1681.7
GTEX-12ZZW-0011-R6a-SM-5DUVZ
50-59 years, male
1646.1
GTEX-1IGQW-0011-R6a-SM-CM2TC
60-69 years, male
1640.8
GTEX-1C6WA-0011-R6a-SM-7PBYP
60-69 years, male
1609.4
GTEX-12WSF-0011-R6b-SM-5HL8W
70-79 years, male
1599.5
GTEX-12WSH-0011-R6b-SM-5P9F4
50-59 years, male
1546.9
GTEX-UTHO-0011-R6A-SM-3GIJW
60-69 years, male
1538.4
GTEX-1EMGI-0011-R6a-SM-9OSVR
60-69 years, male
1524.7
GTEX-X261-0011-R6B-SM-4E3J8
50-59 years, male
1509.3
GTEX-ZUA1-0011-R6b-SM-4YCDD
40-49 years, male
1509.2
GTEX-16XZZ-0011-R6a-SM-7IGNE
60-69 years, male
1502.0
GTEX-1HFI6-0011-R6b-SM-CM2SR
40-49 years, male
1493.4
GTEX-1I1GR-0011-R6a-SM-CMKHI
60-69 years, male
1485.2
GTEX-1IY9M-0011-R6a-SM-D4P28
60-69 years, male
1482.2
GTEX-145MI-0011-R6a-SM-5PNZA
60-69 years, female
1474.7
GTEX-12ZZZ-0011-R6b-SM-5EQ5B
60-69 years, male
1465.3
GTEX-1EN7A-0011-R6b-SM-9QEHW
40-49 years, male
1460.7
GTEX-139TT-0011-R6b-SM-5J2ON
60-69 years, male
1454.5
GTEX-16Z82-0011-R6a-SM-7IGLV
60-69 years, male
1442.2
GTEX-1LVA9-0011-R6a-SM-EYYWH
50-59 years, male
1435.9
GTEX-TSE9-0011-R6A-SM-3DB7L
60-69 years, female
1432.1
GTEX-Z93S-0011-R6b-SM-4RGNJ
50-59 years, male
1429.1
GTEX-YJ89-0011-R6a-SM-4V6GN
60-69 years, male
1425.8
GTEX-X585-0011-R6A-SM-46MVJ
50-59 years, male
1415.0
GTEX-1N2DW-0011-R6a-SM-EZ6MW
50-59 years, male
1409.4
GTEX-1HR9M-0011-R6a-SM-ARU6Z
60-69 years, male
1407.8
GTEX-1EWIQ-0011-R6a-SM-ARU8G
70-79 years, female
1403.0
GTEX-11OF3-0011-R6b-SM-57WBG
60-69 years, male
1391.8
GTEX-13G51-0011-R6b-SM-5LZX4
60-69 years, male
1375.3
GTEX-14E7W-0011-R6b-SM-5ZZV3
70-79 years, male
1375.0
GTEX-11NUK-0011-R6a-SM-DO93O
50-59 years, male
1374.0
GTEX-T2IS-0011-R6A-SM-32QP2
20-29 years, female
1371.4
GTEX-12WSC-0011-R6b-SM-5LU99
50-59 years, male
1367.2
GTEX-1OJC4-0011-R6a-SM-EXOIM
50-59 years, female
1365.5
GTEX-1BAJH-0011-R6b-SM-7MXT2
70-79 years, male
1357.9
GTEX-1AX9I-0011-R6a-SM-7P8PK
60-69 years, male
1354.9
GTEX-144GO-0011-R6b-SM-5S2V7
60-69 years, male
1346.6
GTEX-1E1VI-0011-R6a-SM-9OSVJ
50-59 years, male
1331.4
GTEX-139TS-0011-R6a-SM-5J2OB
60-69 years, male
1316.5
GTEX-11ONC-0011-R6a-SM-DO12A
60-69 years, male
1311.8
GTEX-13NYS-0011-R6b-SM-5MR3R
60-69 years, male
1311.5
GTEX-1A3MX-0011-R6a-SM-7P8S9
60-69 years, male
1310.9
GTEX-1CB4H-0011-R6b-SM-CKZNK
60-69 years, female
1308.8
GTEX-12ZZX-0011-R6a-SM-5HL8L
40-49 years, female
1305.3
GTEX-1LH75-0011-R6a-SM-EXURS
50-59 years, female
1304.5
GTEX-17MF6-0011-R6b-SM-7LG6G
40-49 years, male
1304.0
GTEX-ZF28-0011-R6a-SM-4WKHI
60-69 years, male
1303.1
GTEX-1I1HK-0011-R6b-SM-CM2T9
60-69 years, male
1301.7
GTEX-145MH-0011-R6a-SM-5PNZ7
50-59 years, male
1283.9
GTEX-14C5O-0011-R6a-SM-5YYB6
60-69 years, male
1282.7
GTEX-13FHP-0011-R6b-SM-5LZY6
50-59 years, male
1282.3
GTEX-13PL6-0011-R6a-SM-5O9C5
40-49 years, female
1266.9
GTEX-11NV4-0011-R6a-SM-57WD5
60-69 years, male
1266.7
GTEX-1GMR8-0011-R6b-SM-CE6RC
50-59 years, female
1266.3
GTEX-QVUS-0011-R6A-SM-3GACX
60-69 years, female
1264.0
GTEX-1GZ2Q-0011-R6a-SM-9OSWM
40-49 years, male
1262.5
GTEX-1192X-0011-R6a-SM-DNZZB
50-59 years, male
1258.7
GTEX-14LZ3-0011-R6a-SM-D3L8C
60-69 years, female
1258.6
GTEX-1C64N-0011-R6a-SM-7IGO9
50-59 years, male
1243.4
GTEX-1H3NZ-0011-R6b-SM-A9G1N
60-69 years, male
1242.2
GTEX-13OW5-0011-R6b-SM-5O9BW
60-69 years, male
1241.6
GTEX-13IVO-0011-R6b-SM-5K7X9
70-79 years, male
1240.9
GTEX-13PDP-0011-R6b-SM-5O9D5
60-69 years, male
1238.1
GTEX-11GSP-0011-R6b-SM-57WBQ
60-69 years, female
1234.5
GTEX-11DXY-0011-R6a-SM-DNZZF
60-69 years, male
1231.5
GTEX-11TTK-0011-R6b-SM-DNZZW
60-69 years, female
1230.4
GTEX-1HT8W-0011-R6a-SM-CM2SY
30-39 years, male
1217.0
GTEX-18A6Q-0011-R6a-SM-7LG6R
60-69 years, male
1215.8
GTEX-1H3VE-0011-R6b-SM-A9G1M
40-49 years, male
1208.5
GTEX-1HUB1-0011-R6b-SM-ARU6W
60-69 years, male
1202.7
GTEX-1LB8K-0011-R6a-SM-EXURT
50-59 years, male
1197.8
GTEX-1E2YA-0011-R6b-SM-CE6R2
50-59 years, male
1196.6
GTEX-1ICLZ-0011-R6b-SM-A9G3A
60-69 years, male
1195.4
GTEX-1GF9W-0011-R6b-SM-9QEIE
60-69 years, male
1192.7
GTEX-18464-0011-R6a-SM-731CA
50-59 years, male
1189.9
GTEX-1CB4J-0011-R6b-SM-9WG7B
60-69 years, male
1189.2
GTEX-XMD1-0011-R6A-SM-4AT5K
60-69 years, male
1186.2
GTEX-13SLW-0011-R6a-SM-5O9BH
70-79 years, male
1178.4
GTEX-1IKOH-0011-R6b-SM-A9G1W
60-69 years, female
1170.0
GTEX-18465-0011-R6a-SM-7LG6M
50-59 years, male
1169.9
GTEX-1JMLX-0011-R6b-SM-EXUT9
50-59 years, male
1168.1
GTEX-1B8L1-0011-R6b-SM-7P8SU
60-69 years, male
1163.0
GTEX-17JCI-0011-R6a-SM-72D6D
60-69 years, female
1156.3
GTEX-17HG3-0011-R6b-SM-79OK8
30-39 years, female
1155.5
GTEX-1F75A-0011-R6b-SM-9QEHD
50-59 years, male
1149.7
GTEX-1I19N-0011-R6b-SM-CY8GC
70-79 years, male
1148.3
GTEX-12584-0011-R6a-SM-5NQ7C
40-49 years, male
1146.1
GTEX-YFC4-0011-R6b-SM-4V6E1
40-49 years, female
1144.1
GTEX-1JJEA-0011-R6a-SM-EXUS1
60-69 years, male
1143.0
GTEX-S7SE-0011-R6A-SM-2XCD9
50-59 years, male
1139.8
GTEX-1HGF4-0011-R6a-SM-A96S1
60-69 years, male
1138.3
GTEX-WL46-0011-R6A-SM-3LK6X
50-59 years, male
1137.3
GTEX-13JVG-0011-R6a-SM-5MR3E
60-69 years, male
1133.3
GTEX-1GN2E-0011-R6b-SM-9QEII
60-69 years, male
1132.5
GTEX-ZDXO-0011-R6a-SM-4WWCT
60-69 years, male
1124.3
GTEX-QMR6-0011-R6A-SM-32PKP
50-59 years, male
1122.4
GTEX-1F48J-0011-R6a-SM-CJI2F
50-59 years, female
1119.9
GTEX-14PHW-0011-R6b-SM-5YYAR
60-69 years, female
1118.6
GTEX-13FLV-0011-R6a-SM-5KLZ4
50-59 years, male
1112.7
GTEX-1HBPM-0011-R6b-SM-ARU73
60-69 years, female
1112.2
GTEX-1F75W-0011-R6a-SM-9QEI5
60-69 years, male
1102.1
GTEX-1GF9V-0011-R6b-SM-CJI2Q
60-69 years, male
1101.2
GTEX-1GZ4I-0011-R6b-SM-9WG77
50-59 years, male
1098.9
GTEX-ZVZQ-0011-R6b-SM-57WAY
60-69 years, female
1094.4
GTEX-1B8SF-0011-R6b-SM-7MXUD
60-69 years, male
1091.3
GTEX-1H4P4-0011-R6a-SM-ARZLR
50-59 years, male
1090.3
GTEX-17EVQ-0011-R6a-SM-7189U
70-79 years, male
1087.3
GTEX-1EH9U-0011-R6a-SM-CJI2C
60-69 years, male
1085.9
GTEX-17EVP-0011-R6b-SM-7LT99
40-49 years, male
1083.2
GTEX-QDT8-0011-R6A-SM-32PKI
30-39 years, female
1081.0
GTEX-13X6J-0011-R6a-SM-5PNUB
50-59 years, male
1077.6
GTEX-1F75B-0011-R6a-SM-ARZMI
50-59 years, male
1075.2
GTEX-147GR-0011-R6b-SM-5S2RO
60-69 years, male
1071.0
GTEX-13O3Q-0011-R6a-SM-5KM2V
50-59 years, male
1069.9
GTEX-1I1GQ-0011-R6b-SM-CE6SE
50-59 years, male
1069.4
GTEX-1J8Q2-0011-R6a-SM-CKZPG
60-69 years, male
1060.2
GTEX-1H1ZS-0011-R6b-SM-A9G1F
70-79 years, male
1055.1
GTEX-1MA7X-0011-R6a-SM-EYYWR
60-69 years, male
1053.9
GTEX-13OVJ-0011-R6b-SM-5L3GH
50-59 years, female
1045.5
GTEX-13NYB-0011-R6b-SM-5MR51
40-49 years, male
1037.3
GTEX-13JUV-0011-R6b-SM-5LZYG
40-49 years, female
1036.0
GTEX-ZVT3-0011-R6a-SM-51MSU
60-69 years, female
1033.3
GTEX-1C6VR-0011-R6a-SM-7MXU8
60-69 years, male
1029.3
GTEX-1F6I4-0011-R6a-SM-CE6R4
60-69 years, male
1027.3
GTEX-P44H-0011-R6A-SM-2XCF3
40-49 years, male
1022.6
GTEX-1B933-0011-R6b-SM-79OOQ
50-59 years, female
1022.2
GTEX-14BIM-0011-R6b-SM-5S2VB
60-69 years, female
1020.9
GTEX-1MJK2-0011-R6a-SM-EXUS3
50-59 years, male
1020.4
GTEX-ZXG5-0011-R6a-SM-57WBZ
60-69 years, male
1019.9
GTEX-1A8G6-0011-R6b-SM-7P8PE
60-69 years, male
1019.2
GTEX-1F88E-0011-R6a-SM-CKZNW
60-69 years, female
1019.2
GTEX-145LS-0011-R6b-SM-5PNWL
60-69 years, female
1019.1
GTEX-1I1CD-0011-R6b-SM-CJI3Y
60-69 years, female
1019.0
GTEX-T5JC-0011-R6A-SM-D3L87
20-29 years, male
1018.6
GTEX-13SLX-0011-R6a-SM-5PNX4
50-59 years, female
1018.2
GTEX-183WM-0011-R6b-SM-72D66
70-79 years, female
1017.8
GTEX-1HB9E-0011-R6b-SM-A9G1P
50-59 years, male
1017.8
GTEX-QVJO-0011-R6A-SM-2S1QN
60-69 years, female
1017.4
GTEX-132Q8-0011-R6a-SM-5K7YN
60-69 years, male
1011.9
GTEX-15RJE-0011-R6a-SM-7DHL8
40-49 years, male
1010.5
GTEX-T6MN-0011-R6A-SM-32QP8
50-59 years, male
1008.6
GTEX-ZE7O-0011-R6a-SM-57WCI
50-59 years, female
1003.8
GTEX-11O72-0011-R6a-SM-DNZZQ
40-49 years, male
1003.2
GTEX-14PJM-0011-R6a-SM-6872G
50-59 years, female
1002.1
GTEX-13QJC-0011-R6a-SM-5S2VI
60-69 years, female
997.0
GTEX-1H1DG-0011-R6a-SM-A9SK8
70-79 years, male
994.8
GTEX-1CB4G-0011-R6b-SM-9QEGO
60-69 years, male
990.9
GTEX-ZE9C-0011-R6a-SM-4WWCW
60-69 years, male
988.9
GTEX-131XW-0011-R6b-SM-5K7XY
50-59 years, female
987.1
GTEX-1399T-0011-R6b-SM-5K7YL
60-69 years, male
979.6
GTEX-15EO6-0011-R6a-SM-7KULK
50-59 years, male
973.8
GTEX-OHPN-0011-R6A-SM-2I5FH
60-69 years, female
973.2
GTEX-11WQC-0011-R6a-SM-5BC77
60-69 years, male
972.8
GTEX-1GZHY-0011-R6a-SM-9OSWL
50-59 years, female
972.6
GTEX-1445S-0011-R6b-SM-5PNUF
60-69 years, male
970.9
GTEX-1EX96-0011-R6a-SM-9OSVV
50-59 years, male
968.0
GTEX-1H1CY-0011-R6b-SM-CE6RP
60-69 years, female
967.0
GTEX-N7MS-0011-R6a-SM-2HMJ4
60-69 years, male
962.9
GTEX-ZAB4-0011-R6a-SM-4SOKD
40-49 years, male
956.0
GTEX-NPJ7-0011-R6a-SM-2I3G7
60-69 years, female
955.5
GTEX-13S7M-0011-R6b-SM-5O9D8
60-69 years, female
953.2
GTEX-1R46S-0011-R6a-SM-EZ6N6
60-69 years, female
952.9
GTEX-14BIN-0011-R6a-SM-5S2RH
50-59 years, female
952.6
GTEX-145LU-0011-R6a-SM-5S2V6
50-59 years, male
943.9
GTEX-15G19-0011-R6b-SM-7KUM3
60-69 years, male
941.8
GTEX-1GN1U-0011-R6a-SM-CKZNO
50-59 years, female
939.9
GTEX-1F6IF-0011-R6a-SM-9QEIO
60-69 years, male
935.3
GTEX-17HHY-0011-R6b-SM-7189Z
50-59 years, male
933.5
GTEX-1HBPI-0011-R6b-SM-A9G1T
50-59 years, male
929.2
GTEX-13X6K-0011-R6a-SM-5P9K6
60-69 years, female
928.7
GTEX-XLM4-0011-R6A-SM-4AT4B
60-69 years, male
925.9
GTEX-1F7RK-0011-R6b-SM-ARZMG
20-29 years, female
924.4
GTEX-1GTWX-0011-R6a-SM-A96RN
60-69 years, male
923.2
GTEX-NPJ8-0011-R6a-SM-2HMKB
40-49 years, male
913.7
GTEX-R55E-0011-R6A-SM-2TC5T
20-29 years, male
913.4
GTEX-131XH-0011-R6a-SM-5KM2G
50-59 years, male
913.0
GTEX-13OVH-0011-R6b-SM-5LUAX
70-79 years, male
910.9
GTEX-1IDJH-0011-R6a-SM-A9SLZ
70-79 years, male
906.9
GTEX-P44G-0011-R6A-SM-2XCD3
30-39 years, male
895.6
GTEX-13FLW-0011-R6b-SM-5L3EN
50-59 years, male
895.5
GTEX-RU72-0011-R6A-SM-2TF71
50-59 years, female
895.4
GTEX-17F97-0011-R6a-SM-7EPGS
50-59 years, male
891.8
GTEX-WZTO-0011-R6B-SM-4E3J6
40-49 years, male
891.6
GTEX-14PQA-0011-R6b-SM-6AJAM
50-59 years, female
888.0
GTEX-15ER7-0011-R6a-SM-6M46M
20-29 years, female
886.2
GTEX-13OW6-0011-R6b-SM-5L3GR
50-59 years, male
884.8
GTEX-13RTJ-0011-R6a-SM-5SI74
60-69 years, male
877.6
GTEX-Y8DK-0011-R6A-SM-4V6EG
60-69 years, male
876.5
GTEX-13OVL-0011-R6a-SM-5L3G4
50-59 years, male
871.5
GTEX-13N1W-0011-R6a-SM-5MR43
70-79 years, male
864.4
GTEX-1H23P-0011-R6b-SM-A9G1H
60-69 years, male
857.3
GTEX-WWYW-0011-R6A-SM-3NB3G
50-59 years, female
853.2
GTEX-13VXU-0011-R6a-SM-5SI71
50-59 years, male
851.6
GTEX-14ASI-0011-R6a-SM-5S2UZ
60-69 years, male
849.5
GTEX-14JG1-0011-R6b-SM-5YYAU
40-49 years, male
842.5
GTEX-S7PM-0011-R6A-SM-3NM8F
60-69 years, male
841.4
GTEX-12ZZY-0011-R6b-SM-5EGLA
60-69 years, male
841.0
GTEX-NL3H-0011-R6a-SM-2I3G8
60-69 years, male
840.9
GTEX-PWO3-0011-R6A-SM-2I5F3
60-69 years, female
838.1
GTEX-1GMRU-0011-R6b-SM-A9G3B
50-59 years, male
835.7
GTEX-16YQH-0011-R6b-SM-7938T
30-39 years, male
834.9
GTEX-14JIY-0011-R6b-SM-68714
60-69 years, male
828.6
GTEX-NL4W-0011-R6a-SM-2I3GA
50-59 years, male
827.5
GTEX-13OW8-0011-R6b-SM-5L3I3
60-69 years, male
823.1
GTEX-1IKK5-0011-R6b-SM-A9SLY
60-69 years, male
823.1
GTEX-12WSE-0011-R6b-SM-5LZV5
20-29 years, male
818.8
GTEX-N7MT-0011-R6a-SM-2I3G3
60-69 years, female
817.3
GTEX-13O3O-0011-R6b-SM-5P9GY
60-69 years, female
816.4
GTEX-WVLH-0011-R6A-SM-3MJFZ
50-59 years, male
815.6
GTEX-12WSM-0011-R6b-SM-5EQ6N
60-69 years, male
813.8
GTEX-14BMV-0011-R6b-SM-5YY9I
50-59 years, male
802.1
GTEX-1313W-0011-R6a-SM-5K7XO
50-59 years, female
793.2
GTEX-13112-0011-R6b-SM-5DUVK
50-59 years, male
787.2
GTEX-1J1OQ-0011-R6a-SM-ARU86
40-49 years, female
783.6
GTEX-13N2G-0011-R6b-SM-5MR3P
60-69 years, male
781.3
GTEX-1IL2U-0011-R6a-SM-ARZLG
60-69 years, male
779.2
GTEX-15CHQ-0011-R6b-SM-69LOX
60-69 years, male
768.6
GTEX-14ABY-0011-R6a-SM-5PNWR
50-59 years, male
749.7
GTEX-X4XX-0011-R6B-SM-46MWP
60-69 years, male
744.7
GTEX-1GN73-0011-R6b-SM-CJI2T
60-69 years, male
726.0
GTEX-13FXS-0011-R6b-SM-5L3F7
50-59 years, male
725.2
GTEX-Q2AG-0011-R6A-SM-2HML7
40-49 years, female
719.7
GTEX-1I1GV-0011-R6b-SM-ARU6Q
60-69 years, male
718.1
GTEX-15DYW-0011-R6b-SM-6PALA
60-69 years, female
703.7
GTEX-11DXW-0011-R6a-SM-DO11N
40-49 years, male
692.4
GTEX-15DCD-0011-R6b-SM-6M46L
70-79 years, female
689.0
GTEX-WHSE-0011-R6A-SM-3P5ZP
20-29 years, male
687.0
GTEX-144GL-0011-R6a-SM-5P9HW
50-59 years, male
686.6
GTEX-1I1GS-0011-R6b-SM-CMKHE
60-69 years, male
678.5
GTEX-1B996-0011-R6b-SM-793AQ
60-69 years, male
672.4
GTEX-14PN4-0011-R6b-SM-69LQU
50-59 years, male
629.7
GTEX-X4EP-0011-R6A-SM-3P629
60-69 years, female
498.6
GTEX-12WSD-0011-R6b-SM-5P9JU
60-69 years, female
413.0
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NUCLEUS ACCUMBENS - FANTOM5 CAGEi
RNA expression in human tissues obtained through Cap Analysis of Gene Expression (CAGE) generated by the FANTOM5 project are reported as Scaled Tags Per Million.